/* Comprehensive record and data-entry pages for LamiOmicsDB.
   This file deliberately reuses the approved homepage tokens and global shell.
   It is loaded only by the new/expanded scientific pages, never by the homepage. */

.complete-record-page { overflow: clip; }
.complete-record-content { padding-top: 26px; }
.complete-width { width: 100%; }
.missing-value,
.missing-prose { color: var(--muted-faint); }
.numeric { text-align: right; font-variant-numeric: tabular-nums; }
.coordinate { font-family: var(--font-mono); font-variant-numeric: tabular-nums; white-space: nowrap; }
.scientific-name { color: var(--ink); font-family: var(--font-editorial); }
.scientific-name:hover { color: var(--gold-800); }
.accession-link,
.gene-symbol-link { color: var(--lavender-700); font-weight: 700; text-decoration: none; }
.accession-link:hover,
.gene-symbol-link:hover { color: var(--gold-800); text-decoration: underline; text-underline-offset: 3px; }
.row-actions,
.section-actions { display: flex; align-items: center; flex-wrap: wrap; gap: 7px; }
.result-count,
.catalogue-result-count { color: var(--muted); font-size: 13px; white-space: nowrap; }
.result-count strong,
.catalogue-result-count strong { color: var(--ink); font-size: 15px; font-variant-numeric: tabular-nums; }

/* Shared identity areas */
.species-record-identity,
.gene-record-identity,
.research-record-heading,
.record-identity.accession-record-identity {
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto;
  align-items: end;
  gap: 30px;
  padding: 24px 0 28px;
  border-bottom: 1px solid var(--line-strong);
}
.species-record-copy,
.gene-record-title { min-width: 0; }
.taxonomy-line { display: flex; align-items: center; flex-wrap: wrap; gap: 7px; margin-bottom: 6px; color: var(--muted); font-size: 13px; }
.taxonomy-line a { color: var(--lavender-700); font-weight: 600; }
.species-record-identity h1,
.gene-record-identity h1,
.research-record-heading h1 {
  margin: 0;
  color: var(--ink);
  font-family: var(--font-editorial);
  font-size: clamp(40px, 4vw, 62px);
  font-weight: 500;
  line-height: 1.02;
  letter-spacing: -.045em;
  overflow-wrap: anywhere;
}
.research-record-heading h1 { font-size: clamp(34px, 3.2vw, 50px); }
.research-record-heading h1 span { color: var(--gold-700); font-size: .62em; font-style: normal; letter-spacing: -.02em; }
.gene-record-identity h1 { font-family: var(--font-mono); font-size: clamp(43px, 4.6vw, 68px); letter-spacing: -.04em; }
.species-name-line,
.gene-identity-meta,
.research-record-meta { display: flex; align-items: center; flex-wrap: wrap; gap: 7px 17px; margin-top: 9px; color: var(--muted); }
.species-name-line strong { color: var(--ink); font-size: 17px; }
.alias-summary { overflow-wrap: anywhere; }
.alias-summary > span { color: var(--ink-soft); }
.species-record-lead { max-width: 860px; margin: 14px 0 0; color: var(--muted); font-size: 16px; line-height: 1.62; }
.species-record-actions { display: flex; align-items: center; justify-content: flex-end; flex-wrap: wrap; gap: 8px; }
.gene-full-name { max-width: 820px; margin: 7px 0 0; color: var(--ink-soft); font-family: var(--font-editorial); font-size: clamp(18px, 2vw, 24px); line-height: 1.35; }
.gene-identity-meta span { display: grid; gap: 2px; }
.gene-identity-meta strong { color: var(--muted-faint); font-size: 10px; letter-spacing: .08em; text-transform: uppercase; }
.research-record-meta code,
.research-record-meta .accession-link { max-width: 320px; overflow: hidden; text-overflow: ellipsis; white-space: nowrap; }

.gene-evidence-strip,
.record-meta-strip,
.expression-summary-strip {
  display: grid;
  grid-template-columns: repeat(5, minmax(0, 1fr));
  background: var(--surface-raised);
  border: 1px solid var(--line-strong);
  border-top: 0;
}
.gene-evidence-strip > div,
.record-meta-strip > div,
.expression-summary-strip > div { min-width: 0; padding: 13px 16px; border-right: 1px solid var(--line); }
.gene-evidence-strip > div:last-child,
.record-meta-strip > div:last-child,
.expression-summary-strip > div:last-child { border-right: 0; }
.gene-evidence-strip span,
.record-meta-strip span,
.expression-summary-strip span { display: block; color: var(--muted); font-size: 11px; }
.gene-evidence-strip strong,
.record-meta-strip strong,
.expression-summary-strip strong { display: block; margin-top: 2px; color: var(--ink); font-size: 18px; font-variant-numeric: tabular-nums; overflow-wrap: anywhere; }
.status-inline.status-available { color: var(--status-available); }
.status-inline.status-unavailable { color: var(--status-unavailable); }

/* Species evidence ribbon */
.evidence-ribbon {
  display: grid;
  grid-template-columns: minmax(230px, .72fr) minmax(0, 2fr);
  gap: 18px;
  padding: 18px 20px;
  background: linear-gradient(120deg, color-mix(in srgb, var(--lavender-75) 80%, var(--surface)), var(--surface));
  border: 1px solid var(--line-strong);
}
.evidence-ribbon-intro h2 { margin: 1px 0 3px; font-family: var(--font-editorial); font-size: 20px; }
.evidence-ribbon-intro p { margin: 0; color: var(--muted); font-size: 12px; }
.evidence-ribbon-grid { display: grid; grid-template-columns: repeat(4, minmax(0, 1fr)); border-left: 1px solid var(--line); }
.evidence-layer-cell { min-width: 0; display: grid; grid-template-columns: 38px minmax(0, 1fr) 21px; align-items: center; gap: 8px; padding: 10px 12px; border-right: 1px solid var(--line); color: var(--ink); }
.evidence-layer-cell:hover { background: var(--surface-raised); }
.evidence-layer-icon { width: 34px; height: 34px; display: grid; place-items: center; color: var(--lavender-700); }
.evidence-layer-icon svg { width: 31px; height: 31px; }
.evidence-layer-cell strong,
.evidence-layer-cell small { display: block; }
.evidence-layer-cell strong { font-family: var(--font-editorial); font-size: 13px; }
.evidence-layer-cell small { color: var(--muted); font-size: 10px; }
.evidence-layer-cell .status-shape { width: 20px; height: 20px; }
.evidence-layer-cell.is-available .status-shape { color: var(--status-available); }
.evidence-layer-cell.is-metadata-only .status-shape { color: var(--status-metadata); }
.evidence-layer-cell.is-unavailable .status-shape { color: var(--status-unavailable); }

/* Main record layout */
.record-layout { display: grid; grid-template-columns: minmax(0, 1fr) 300px; align-items: start; gap: 28px; }
.record-layout-species { margin-top: 28px; }
.record-main { min-width: 0; }
.record-aside-unified,
.genome-side-ledger { min-width: 0; display: grid; align-content: start; gap: 16px; }
.record-aside-unified { position: sticky; top: calc(var(--header-primary-height) + var(--header-nav-height) + 64px); }
.section-block { scroll-margin-top: calc(var(--header-primary-height) + var(--header-nav-height) + 62px); }
.section-block + .section-block { margin-top: 42px; }
.section-heading { display: flex; align-items: end; justify-content: space-between; gap: 22px; margin-bottom: 16px; padding-bottom: 10px; border-bottom: 1px solid var(--line-strong); }
.section-heading h2 { margin: 0; color: var(--ink); font-family: var(--font-editorial); font-size: 25px; font-weight: 600; }
.section-heading p { max-width: 720px; margin: 4px 0 0; color: var(--muted); font-size: 13px; line-height: 1.48; }
.record-intro { max-width: 880px; margin: 0; color: var(--ink-soft); font-family: var(--font-editorial); font-size: 18px; line-height: 1.72; }
.species-overview-grid { display: grid; grid-template-columns: minmax(0, 1.25fr) minmax(260px, .75fr); gap: 28px; }
.species-portrait-panel { margin: 0; min-height: 390px; display: grid; grid-template-rows: 1fr auto; background: linear-gradient(145deg, var(--surface-soft), var(--surface-raised)); border: 1px solid var(--line); overflow: hidden; }
.species-portrait-panel img { width: 100%; height: 100%; min-height: 350px; object-fit: contain; padding: 18px; }
.species-portrait-panel figcaption { padding: 10px 13px; color: var(--muted); border-top: 1px solid var(--line); font-size: 11px; }
.editorial-note-list { margin-top: 22px; border-top: 1px solid var(--line); }
.editorial-note-list article { display: grid; grid-template-columns: 170px minmax(0, 1fr); gap: 20px; padding: 15px 0; border-bottom: 1px solid var(--line); }
.editorial-note-list h3 { margin: 0; font-family: var(--font-editorial); font-size: 16px; }
.editorial-note-list p { margin: 0; color: var(--muted); line-height: 1.58; }
.record-prose-facts { margin: 22px 0 0; }
.record-prose-facts div { display: grid; grid-template-columns: 170px minmax(0, 1fr); gap: 20px; padding: 13px 0; border-top: 1px solid var(--line); }
.record-prose-facts dt { color: var(--ink); font-family: var(--font-editorial); font-weight: 600; }
.record-prose-facts dd { margin: 0; color: var(--muted); line-height: 1.55; }
.panel { overflow: hidden; }
.panel-header h2 { font-size: 17px; }
.record-fact-list { margin: 0; }
.record-fact-list > div { display: grid; grid-template-columns: minmax(95px, .8fr) minmax(0, 1.2fr); gap: 12px; padding: 9px 0; border-bottom: 1px solid var(--line); }
.record-fact-list > div:last-child { border-bottom: 0; }
.record-fact-list dt { color: var(--muted); font-size: 12px; }
.record-fact-list dd { min-width: 0; margin: 0; color: var(--ink); font-weight: 600; text-align: right; overflow-wrap: anywhere; }
.record-fact-list dd.mono { font-size: 12px; }
.tree-membership-list { display: grid; }
.tree-membership-list a { display: flex; align-items: center; justify-content: space-between; gap: 10px; padding: 9px 0; border-bottom: 1px solid var(--line); }
.tree-membership-list a:last-child { border-bottom: 0; }
.tree-membership-list strong { color: var(--ink-soft); font-size: 12px; }
.layer-name-cell { display: flex; align-items: center; gap: 8px; }
.layer-name-cell svg { width: 24px; height: 24px; color: var(--lavender-700); }
.identity-cell { min-width: 0; display: grid; gap: 2px; }
.identity-cell strong { color: var(--ink); }
.identity-cell small { color: var(--muted); line-height: 1.35; }
.sequence-record-list { border-top: 1px solid var(--line); }
.sequence-record-row { display: grid; grid-template-columns: minmax(230px, 1.3fr) minmax(150px, .7fr) 90px auto; align-items: center; gap: 14px; padding: 11px 0; border-bottom: 1px solid var(--line); }
.sequence-record-row > div { display: grid; }
.sequence-record-row code { overflow: hidden; text-overflow: ellipsis; }
.pathway-record-summary { display: grid; grid-template-columns: minmax(0, 1fr) auto; align-items: center; gap: 20px; padding: 17px 19px; background: var(--surface-soft); border: 1px solid var(--line); }
.pathway-record-summary h3 { margin: 0; font-family: var(--font-editorial); }
.pathway-record-summary p { margin: 5px 0 0; color: var(--muted); }

/* Omics overview */
.omics-ledger { border-top: 1px solid var(--line-strong); }
.omics-ledger-row { display: grid; grid-template-columns: minmax(260px, 1.15fr) minmax(200px, .7fr) minmax(360px, 1.3fr) auto; align-items: center; gap: 20px; padding: 18px 0; border-bottom: 1px solid var(--line-strong); }
.omics-ledger-identity { display: grid; grid-template-columns: 50px minmax(0, 1fr); gap: 13px; }
.omics-ledger-icon { width: 48px; height: 48px; display: grid; place-items: center; color: var(--lavender-700); }
.omics-ledger-icon svg { width: 44px; height: 44px; }
.omics-ledger-identity h3 { margin: 0; font-family: var(--font-editorial); font-size: 20px; }
.omics-ledger-identity p,
.omics-ledger-status small { margin: 4px 0 0; color: var(--muted); font-size: 12px; line-height: 1.42; }
.omics-ledger-status { display: grid; justify-items: start; gap: 4px; }
.omics-ledger-facts { display: grid; grid-template-columns: repeat(5, minmax(0, 1fr)); margin: 0; }
.omics-ledger-facts div { min-width: 0; padding: 3px 10px; border-left: 1px solid var(--line); }
.omics-ledger-facts dt { color: var(--muted); font-size: 9px; text-transform: uppercase; letter-spacing: .05em; }
.omics-ledger-facts dd { margin: 3px 0 0; color: var(--ink); font-size: 12px; font-weight: 600; overflow-wrap: anywhere; }
.cross-layer-grid,
.record-stat-grid,
.assembly-summary-grid { display: grid; grid-template-columns: repeat(4, minmax(0, 1fr)); border: 1px solid var(--line-strong); }
.cross-layer-stat,
.record-stat-grid > div,
.assembly-metric { padding: 20px; border-right: 1px solid var(--line); }
.cross-layer-stat:last-child,
.record-stat-grid > div:last-child,
.assembly-metric:last-child { border-right: 0; }
.cross-layer-stat strong,
.record-stat-grid strong,
.assembly-metric strong { display: block; color: var(--gold-700); font-family: var(--font-editorial); font-size: 29px; font-variant-numeric: tabular-nums; }
.cross-layer-stat span,
.record-stat-grid span,
.assembly-metric span { display: block; margin-top: 3px; color: var(--muted); font-size: 12px; }
.research-path-list { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.research-path-list > a { min-width: 0; display: grid; grid-template-columns: 42px minmax(0, 1fr) 20px; align-items: center; gap: 12px; padding: 17px; color: var(--ink); border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.research-path-list > a:hover { background: var(--surface-soft); }
.research-path-list > a > svg:first-child { width: 37px; height: 37px; color: var(--lavender-700); }
.research-path-list > a > svg:last-child { width: 18px; height: 18px; color: var(--gold-700); }
.research-path-list strong,
.research-path-list span { display: block; }
.research-path-list strong { font-family: var(--font-editorial); }
.research-path-list span { margin-top: 3px; color: var(--muted); font-size: 12px; }

/* Research toolbar */
.research-toolbar { position: sticky; z-index: 57; width: 100%; min-height: 46px; padding: 0; background: color-mix(in srgb, var(--surface) 96%, transparent); border-top: 1px solid var(--line); border-bottom: 1px solid var(--line-strong); backdrop-filter: blur(16px); }
.research-toolbar-inner { min-height: 46px; display: flex; align-items: center; gap: 10px; }
.research-toolbar-title { min-width: 190px; display: flex; align-items: center; gap: 8px; padding-right: 12px; border-right: 1px solid var(--line); color: var(--ink); font-weight: 700; }
.research-toolbar-title svg { width: 23px; height: 23px; color: var(--lavender-700); }
.research-toolbar-group { display: flex; align-items: center; gap: 5px; }
.research-toolbar-group .button.is-active { color: var(--gold-800); background: var(--gold-75); border-color: var(--gold-500); }
.research-toolbar-search { min-width: 220px; flex: 1 1 340px; height: 36px; display: grid; grid-template-columns: 26px minmax(0, 1fr); align-items: center; padding: 0 10px; background: var(--surface-raised); border: 1px solid var(--line); border-radius: 5px; }
.research-toolbar-search svg { width: 18px; height: 18px; color: var(--muted); }
.research-toolbar-search input { width: 100%; height: 100%; min-width: 0; color: var(--ink); background: transparent; border: 0; outline: 0; }

/* Organelle viewer */
.organelle-viewer-shell { padding-top: 18px; }
.genome-research-stage { background: var(--surface); border: 1px solid var(--line-strong); }
.genome-view-panel { min-height: 640px; }
.genome-view-panel.is-active { display: block; }
.genome-view-panel[data-genome-view="circular"] { display: grid; grid-template-columns: minmax(0, 1fr) 300px; }
.genome-canvas.complete-genome-canvas { min-width: 0; min-height: 690px; display: grid; place-items: center; padding: 14px; overflow: auto; background: radial-gradient(circle at 50% 45%, var(--surface-raised), var(--surface-soft)); }
.complete-genome-canvas .organelle-map-svg { width: min(100%, 1050px); height: auto; transform-origin: center; transition: transform 120ms ease; }
.complete-inspector { min-height: 100%; padding: 20px; background: var(--surface-soft); border-left: 1px solid var(--line-strong); }
.inspector-empty { text-align: center; padding: 50px 10px; }
.inspector-symbol { width: 54px; height: 54px; display: grid; place-items: center; margin: 0 auto 13px; color: var(--gold-700); border: 1px solid var(--line-gold); border-radius: 50%; }
.inspector-symbol svg { width: 29px; height: 29px; }
.inspector-empty h2,
.inspector-selection h2 { margin: 0; font-family: var(--font-editorial); font-size: 22px; }
.inspector-empty p,
.inspector-selection p { color: var(--muted); line-height: 1.5; }
.linear-genome-viewport { width: 100%; min-height: 510px; overflow-x: auto; overflow-y: hidden; background: var(--surface-soft); }
.linear-genome-complete { width: var(--linear-width, 1160px); min-width: 1160px; min-height: 500px; padding: 74px 54px 60px; position: relative; }
.linear-coordinate-ruler { position: relative; height: 46px; margin-bottom: 28px; border-bottom: 1px solid var(--ink-soft); }
.linear-coordinate-ruler::before { content: ""; position: absolute; inset: auto 0 -5px; height: 9px; background: repeating-linear-gradient(90deg, var(--ink-soft) 0 1px, transparent 1px 5%); opacity: .35; }
.linear-coordinate-ruler span { position: absolute; bottom: 12px; color: var(--muted); font-family: var(--font-mono); font-size: 10px; transform: translateX(-50%); }
.linear-coordinate-ruler span:first-child { transform: none; }
.linear-coordinate-ruler span:last-child { transform: translateX(-100%); }
.linear-track-label { display: flex; justify-content: space-between; color: var(--muted); font-size: 11px; }
.linear-feature-track { position: relative; height: 168px; margin: 7px 0; }
.linear-genome-axis { height: 12px; display: grid; place-items: center; }
.linear-genome-axis span { width: 100%; height: 2px; background: var(--ink-soft); }
.linear-feature { position: absolute; left: var(--left); width: max(var(--width), 4px); min-width: 3px; height: 29px; padding: 0; color: #fff; background: var(--feature-color); border: 0; border-radius: 2px; cursor: pointer; overflow: visible; }
.linear-feature::after { content: ""; position: absolute; top: 0; right: -8px; border-top: 14.5px solid transparent; border-bottom: 14.5px solid transparent; border-left: 8px solid var(--feature-color); }
.linear-feature-track.is-forward .linear-feature { bottom: 0; }
.linear-feature span { position: absolute; left: 50%; bottom: calc(36px + (var(--label-lane, 0) * 18px)); max-width: 130px; color: var(--ink-soft); font-family: var(--font-mono); font-size: 9px; line-height: 1; white-space: nowrap; transform: translateX(-2px) rotate(-45deg); transform-origin: left bottom; }
.linear-feature-track.is-reverse .linear-feature { bottom: 0; }
.linear-feature-track.is-reverse .linear-feature::after { right: auto; left: -8px; border-left: 0; border-right: 8px solid var(--feature-color); }
.linear-feature-track.is-reverse .linear-feature span { top: calc(39px + (var(--label-lane, 0) * 18px)); bottom: auto; transform: translateX(-2px) rotate(45deg); transform-origin: left top; }
.linear-feature.is-selected,
.linear-feature:focus-visible { outline: 3px solid var(--focus); outline-offset: 3px; z-index: 4; }
.genome-support-grid { display: grid; grid-template-columns: minmax(0, 1fr) 300px; align-items: start; gap: 26px; margin-top: 30px; }
.feature-table-wrap { max-height: 620px; }
.feature-table-wrap tbody tr.is-selected { background: color-mix(in srgb, var(--focus) 10%, var(--surface)); box-shadow: inset 3px 0 var(--focus); }
.feature-group-swatch { width: 10px; height: 10px; display: inline-block; margin-right: 6px; background: var(--feature-color); border: 1px solid rgba(0,0,0,.12); }
.genome-legend ul { list-style: none; display: grid; gap: 8px; margin: 0; padding: 0; }
.genome-legend li { display: grid; grid-template-columns: 14px minmax(0, 1fr) auto; align-items: center; gap: 8px; }
.genome-legend li > span { width: 12px; height: 12px; background: var(--feature-color); }
.genome-legend li small { color: var(--muted); font-variant-numeric: tabular-nums; }
.source-ledger { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.source-ledger a { min-width: 0; display: grid; grid-template-columns: minmax(90px, .45fr) minmax(0, 1fr) 18px; align-items: center; gap: 10px; padding: 12px; border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); color: var(--ink); }
.source-ledger a:hover { background: var(--surface-soft); }
.source-ledger span { color: var(--muted); font-size: 11px; }
.source-ledger code { overflow: hidden; text-overflow: ellipsis; }
.source-ledger svg { width: 17px; height: 17px; color: var(--gold-700); }

/* Nuclear and files */
.nuclear-record-layout { margin-top: 2px; }
.assembly-summary-grid { margin-bottom: 22px; }
.file-ledger { border-top: 1px solid var(--line); }
.file-ledger article { display: grid; grid-template-columns: 38px minmax(0, 1fr) auto; align-items: center; gap: 12px; padding: 12px 0; border-bottom: 1px solid var(--line); }
.file-ledger article > svg { width: 31px; height: 31px; color: var(--lavender-700); }
.file-ledger article strong,
.file-ledger article small { display: block; }
.file-ledger article small { margin-top: 2px; color: var(--muted); }
.structured-metadata-grid { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); margin: 0; border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.structured-metadata-grid > div { min-width: 0; padding: 13px 15px; border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.structured-metadata-grid dt { color: var(--muted); font-size: 11px; text-transform: uppercase; letter-spacing: .04em; }
.structured-metadata-grid dd { margin: 5px 0 0; color: var(--ink); line-height: 1.45; overflow-wrap: anywhere; }
.editorial-callout { display: grid; grid-template-columns: 24px minmax(0, 1fr); gap: 12px; margin-top: 18px; padding: 15px 17px; background: var(--gold-75); border-left: 3px solid var(--gold-600); }
.editorial-callout svg { width: 22px; height: 22px; color: var(--gold-700); }
.editorial-callout p { margin: 0; color: var(--ink-soft); line-height: 1.55; }

/* Transcriptome */
.expression-summary-strip { grid-template-columns: repeat(4, minmax(0, 1fr)); margin-bottom: 16px; }
.all-nd-alert { margin-bottom: 16px; }
.expression-viewer-stage { display: grid; grid-template-columns: minmax(0, 1fr) 270px; background: var(--surface); border: 1px solid var(--line-strong); }
.expression-legend-complete { grid-column: 1 / -1; min-height: 53px; display: flex; align-items: center; gap: 25px; padding: 9px 16px; border-bottom: 1px solid var(--line); }
.expression-legend-complete > div:first-child { display: grid; }
.expression-legend-complete small { color: var(--muted); }
.sequential-legend { display: flex; align-items: center; gap: 8px; color: var(--muted); font-size: 11px; }
.sequential-legend i { width: 210px; height: 13px; background: linear-gradient(90deg,#f7fafa,#e3eef0,#c6dce1,#9fc2ca,#76a3ae,#527e89,#354f59); border: 1px solid var(--line); }
.nd-key { display: flex; align-items: center; gap: 8px; color: var(--muted); font-size: 11px; }
.nd-key i { width: 20px; height: 15px; background: repeating-linear-gradient(135deg,#f4f0ec 0 3px,#7a7069 3px 4px,#f4f0ec 4px 7px); border: 1px solid #cfc4bb; }
.expression-matrix-viewport { min-width: 0; max-height: 650px; overflow: auto; }
.expression-matrix-complete { min-width: calc(190px + (var(--heatmap-columns) * 76px)); display: grid; grid-template-columns: 190px repeat(var(--heatmap-columns), minmax(76px, 1fr)); }
.heatmap-corner,
.heatmap-column-label,
.heatmap-row-label { position: sticky; z-index: 3; background: var(--surface-soft); border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.heatmap-corner { top: 0; left: 0; z-index: 6; padding: 12px; color: var(--muted); font-size: 11px; }
.heatmap-column-label { top: 0; min-height: 116px; display: flex; align-items: flex-end; justify-content: center; padding: 10px 8px 16px; overflow: visible; }
.heatmap-column-label span { color: var(--muted); font-size: 10px; line-height: 1; white-space: nowrap; transform: translateX(42%) rotate(-45deg); transform-origin: left bottom; }
.heatmap-row-label { left: 0; min-height: 45px; display: flex; align-items: center; padding: 0 12px; }
.heatmap-cell-complete { min-height: 45px; display: grid; place-items: center; padding: 0; color: color-mix(in srgb, #172022 calc(100% - var(--intensity)), #fff var(--intensity)); background: color-mix(in srgb, #f7fafa calc(100% - var(--intensity)), #354f59 var(--intensity)); border: 0; border-right: 1px solid rgba(255,255,255,.32); border-bottom: 1px solid rgba(255,255,255,.32); cursor: pointer; font-family: var(--font-mono); font-size: 10px; }
.heatmap-cell-complete:hover,
.heatmap-cell-complete.is-selected { outline: 3px solid var(--focus); outline-offset: -3px; z-index: 2; }
.heatmap-cell-complete.is-nd { color: #6b5a4f; background: repeating-linear-gradient(135deg,#f4f0ec 0 4px,#7a7069 4px 5px,#f4f0ec 5px 9px); }
.heatmap-inspector-complete { padding: 19px; background: var(--surface-soft); border-left: 1px solid var(--line-strong); }
.heatmap-inspector-complete h2 { margin: 0 0 14px; font-family: var(--font-editorial); font-size: 20px; }
.heatmap-inspector-complete > p:last-child { color: var(--muted); font-size: 12px; line-height: 1.5; }
.nd-table-cell { background: repeating-linear-gradient(135deg,var(--surface-soft) 0 5px,color-mix(in srgb,var(--status-nd) 35%,var(--surface-soft)) 5px 6px,var(--surface-soft) 6px 11px); color: var(--status-nd); font-weight: 700; }
.transcriptome-support-layout { margin-top: 34px; }

/* Scientific states */
.scientific-state-page { min-height: 480px; display: grid; place-items: center; padding: 50px 0; }
.scientific-state { width: min(680px, 100%); display: grid; justify-items: center; text-align: center; padding: 34px; background: var(--surface-raised); border: 1px solid var(--line-strong); }
.scientific-state > svg { width: 54px; height: 54px; margin-bottom: 12px; }
.scientific-state h2,
.scientific-state h3 { margin: 0; font-family: var(--font-editorial); }
.scientific-state h2 { font-size: 28px; }
.scientific-state p { max-width: 580px; color: var(--muted); line-height: 1.58; }
.scientific-state.state-available > svg { color: var(--status-available); }
.scientific-state.state-metadata-only > svg,
.scientific-state.state-metadata > svg { color: var(--status-metadata); }
.scientific-state.state-unavailable > svg { color: var(--status-unavailable); }
.compact-state { min-height: 190px; padding: 26px; }
.state-actions { display: flex; flex-wrap: wrap; justify-content: center; gap: 8px; }
.complete-empty { min-height: 280px; }

/* Catalogue filters and tables */
.catalogue-toolbar { min-height: 55px; display: flex; align-items: center; flex-wrap: wrap; gap: 10px; margin-bottom: 14px; }
.catalogue-search { flex: 1 1 520px; height: 42px; display: grid; grid-template-columns: 28px minmax(0, 1fr) auto; align-items: center; padding-left: 10px; background: var(--surface-raised); border: 1px solid var(--line-strong); border-radius: 5px; }
.catalogue-search > svg { width: 18px; height: 18px; color: var(--muted); }
.catalogue-search input { width: 100%; height: 100%; min-width: 0; color: var(--ink); background: transparent; border: 0; outline: 0; }
.catalogue-search .button { border-top: 0; border-right: 0; border-bottom: 0; border-radius: 0; }
.compact-select { min-width: 190px; display: grid; grid-template-columns: auto minmax(110px, 1fr); align-items: center; gap: 8px; }
.compact-select span { color: var(--muted); font-size: 12px; }
.compact-select select,
.catalogue-filter-grid select { min-height: 42px; color: var(--ink); background: var(--surface-raised); border: 1px solid var(--line-strong); border-radius: 5px; padding: 0 34px 0 10px; }
.catalogue-filter-grid { display: grid; grid-template-columns: minmax(340px, 1fr) repeat(3, minmax(160px, .45fr)) auto auto; align-items: end; gap: 10px; margin-bottom: 12px; }
.catalogue-filter-grid > label { min-width: 0; display: grid; gap: 5px; color: var(--muted); font-size: 11px; }
.filter-search { height: 42px; display: grid; grid-template-columns: 28px minmax(0, 1fr); align-items: center; padding: 0 10px; background: var(--surface-raised); border: 1px solid var(--line-strong); border-radius: 5px; }
.filter-search svg { width: 18px; height: 18px; color: var(--muted); }
.filter-search input { width: 100%; height: 100%; color: var(--ink); background: transparent; border: 0; outline: 0; }
.catalogue-summary-row { display: flex; justify-content: space-between; gap: 16px; margin: 11px 0; color: var(--muted); font-size: 13px; }
.catalogue-summary-row strong { color: var(--ink); }
.comfortable-table tbody td { min-height: 52px; padding-top: 11px; padding-bottom: 11px; }
.compact-table th,
.compact-table td { padding-top: 8px; padding-bottom: 8px; font-size: 12.5px; }
.mini-data-label { display: inline-block; margin: 1px 3px 1px 0; padding: 2px 5px; color: var(--ink-soft); background: var(--surface-soft); border: 1px solid var(--line); border-radius: 2px; font-size: 10px; }
.status-metadata-only { color: var(--status-metadata); }
.status-reference { color: var(--status-reference); }
.table-note { margin: 10px 0 0; color: var(--muted); font-size: 12px; }
.pagination { min-height: 52px; display: flex; align-items: center; justify-content: center; gap: 15px; margin-top: 14px; color: var(--muted); }
.pagination a { min-height: 36px; display: inline-flex; align-items: center; padding: 0 12px; color: var(--ink); background: var(--surface-raised); border: 1px solid var(--line-strong); border-radius: 4px; }
.coverage-inline { min-width: 110px; display: grid; grid-template-columns: 1fr auto; align-items: center; gap: 8px; }
.coverage-inline::before { content: ""; grid-column: 1; grid-row: 1; height: 7px; background: var(--surface-soft); border: 1px solid var(--line); }
.coverage-inline i { grid-column: 1; grid-row: 1; z-index: 1; height: 7px; background: linear-gradient(90deg,var(--lavender-500),var(--gold-600)); }
.coverage-inline strong { grid-column: 2; font-size: 11px; }

/* Gene record */
.gene-record-layout { grid-template-columns: minmax(0, 1fr) 310px; }
.gene-metadata-grid { margin-top: 22px; }
.gene-coordinate-table-wrap { max-height: 640px; }
.gene-sequence-ledger { border-top: 1px solid var(--line); }
.gene-sequence-ledger article { display: grid; grid-template-columns: minmax(250px, .9fr) minmax(300px, 1.35fr) auto; align-items: center; gap: 18px; padding: 13px 0; border-bottom: 1px solid var(--line); }
.sequence-ledger-identity { min-width: 0; display: grid; gap: 2px; }
.sequence-ledger-identity strong { color: var(--ink); }
.sequence-ledger-identity small { color: var(--muted); }
.gene-sequence-ledger pre { min-width: 0; margin: 0; padding: 8px 10px; overflow: hidden; color: var(--ink-soft); background: var(--surface-soft); border: 1px solid var(--line); font-family: var(--font-mono); font-size: 10px; white-space: nowrap; text-overflow: ellipsis; }
.accession-chip-ledger { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.accession-chip-ledger a { min-width: 0; display: grid; grid-template-columns: minmax(0, 1fr) auto 18px; align-items: center; gap: 8px; padding: 12px; color: var(--ink); border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.accession-chip-ledger code { overflow: hidden; text-overflow: ellipsis; }
.accession-chip-ledger span { color: var(--muted); font-size: 10px; }
.accession-chip-ledger svg { width: 16px; height: 16px; color: var(--gold-700); }
.related-gene-list { border-top: 1px solid var(--line); }
.related-gene-list a { display: grid; grid-template-columns: 90px minmax(200px, .8fr) minmax(220px, 1fr) 18px; align-items: center; gap: 12px; padding: 11px 0; color: var(--ink); border-bottom: 1px solid var(--line); }
.related-gene-list span { color: var(--ink-soft); }
.related-gene-list small { color: var(--muted); }
.related-gene-list svg { width: 17px; height: 17px; color: var(--gold-700); }

/* Global omics, taxonomy and accession */
.omics-layer-ledger { border-top: 1px solid var(--line-strong); }
.omics-layer-entry { display: grid; grid-template-columns: 60px minmax(250px, 1fr) minmax(220px, .7fr) 22px; align-items: center; gap: 16px; padding: 17px 0; color: var(--ink); border-bottom: 1px solid var(--line-strong); }
.omics-layer-entry:hover { background: linear-gradient(90deg,transparent,var(--surface-soft),transparent); }
.omics-layer-entry-icon { width: 54px; height: 54px; display: grid; place-items: center; color: var(--lavender-700); }
.omics-layer-entry-icon svg { width: 48px; height: 48px; }
.omics-layer-entry h3 { margin: 0; font-family: var(--font-editorial); font-size: 21px; }
.omics-layer-entry p { margin: 4px 0 0; color: var(--muted); }
.coverage-meter { display: grid; grid-template-columns: minmax(0, 1fr) 42px; align-items: center; gap: 9px; }
.coverage-meter::before { content: ""; grid-column: 1; grid-row: 1; height: 9px; background: var(--surface-soft); border: 1px solid var(--line); }
.coverage-meter i { grid-column: 1; grid-row: 1; z-index: 1; height: 9px; background: linear-gradient(90deg,var(--lavender-500),var(--gold-600)); }
.coverage-meter span { grid-column: 2; color: var(--ink-soft); font-family: var(--font-mono); font-size: 11px; }
.ledger-arrow { width: 20px; height: 20px; color: var(--gold-700); }
.matrix-cell-link { display: grid; grid-template-columns: auto auto; gap: 2px 8px; color: var(--ink); }
.matrix-cell-link strong { font-size: 16px; }
.matrix-cell-link span { color: var(--gold-700); font-size: 11px; }
.matrix-cell-link small { grid-column: 1 / -1; color: var(--muted); font-size: 9px; }
.evidence-definition-row { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 14px; }
.organelle-layer-split { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.organelle-layer-entry { min-height: 230px; display: grid; grid-template-columns: 150px minmax(0, 1fr) 22px; align-items: center; gap: 22px; padding: 24px; color: var(--ink); border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.organelle-layer-entry:hover { background: var(--surface-soft); }
.organelle-layer-art { position: relative; width: 138px; height: 138px; display: grid; place-items: center; }
.organelle-layer-art::before,
.organelle-layer-art::after,
.organelle-layer-art i { content: ""; position: absolute; border-radius: 50%; border: 1px solid var(--lavender-300); }
.organelle-layer-art::before { inset: 2px; }
.organelle-layer-art::after { inset: 18px; border-style: dashed; border-color: var(--gold-500); }
.organelle-layer-art i { inset: 34px; border-width: 7px; border-color: color-mix(in srgb,var(--lavender-500) 55%,transparent); }
.organelle-layer-art svg { width: 48px; height: 48px; color: var(--lavender-700); }
.organelle-layer-entry h2 { margin: 0; font-family: var(--font-editorial); }
.organelle-layer-entry p { color: var(--muted); }
.taxonomy-ledger { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.taxonomy-ledger > a { min-width: 0; display: grid; grid-template-columns: 46px minmax(0, 1fr) 18px; align-items: center; gap: 12px; padding: 15px; color: var(--ink); border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.taxonomy-ledger > a:hover { background: var(--surface-soft); }
.taxonomy-mark { width: 42px; height: 42px; display: grid; place-items: center; color: var(--lavender-700); }
.taxonomy-mark svg { width: 36px; height: 36px; }
.taxonomy-ledger h3 { margin: 0; font-family: var(--font-editorial); }
.taxonomy-ledger p { margin: 3px 0 0; color: var(--muted); font-size: 12px; }
.taxonomy-ledger > a > svg { width: 17px; height: 17px; color: var(--gold-700); }
.genus-inline-index { display: flex; flex-wrap: wrap; gap: 8px; }
.genus-inline-index a { min-height: 39px; display: inline-flex; align-items: center; gap: 9px; padding: 0 12px; color: var(--ink); background: var(--surface-raised); border: 1px solid var(--line); border-radius: 3px; }
.genus-inline-index span { color: var(--muted); font-size: 11px; }
.reference-ledger { border-top: 1px solid var(--line); }
.reference-ledger article { display: grid; grid-template-columns: 55px minmax(0, 1fr) auto; gap: 15px; padding: 14px 0; border-bottom: 1px solid var(--line); }
.reference-year { color: var(--gold-700); font-family: var(--font-mono); font-weight: 700; }
.reference-ledger h3 { margin: 0; font-family: var(--font-editorial); font-size: 16px; }
.reference-ledger p { margin: 4px 0 0; color: var(--muted); font-size: 12px; }
.accession-filters { grid-template-columns: minmax(360px, 1fr) minmax(190px, .35fr) auto auto; }

/* Dark-mode refinements for embedded scientific SVG and heatmaps */
html[data-theme="dark"] .complete-genome-canvas .organelle-map-svg { filter: brightness(.88) saturate(.9); }
html[data-theme="dark"] .heatmap-cell-complete { color: #f7f2fb; }
html[data-theme="dark"] .heatmap-cell-complete.is-nd { color: #ead8c8; background: repeating-linear-gradient(135deg,#201b1a 0 4px,#8d7767 4px 5px,#201b1a 5px 9px); }
html[data-theme="dark"] .linear-feature span { color: var(--ink-soft); text-shadow: 0 1px 1px var(--page); }

/* Responsive */
@media (max-width: 1180px) {
  .record-layout,
  .gene-record-layout { grid-template-columns: minmax(0, 1fr) 270px; gap: 20px; }
  .evidence-ribbon { grid-template-columns: 1fr; }
  .omics-ledger-row { grid-template-columns: minmax(240px, 1fr) minmax(180px, .65fr) minmax(0, 1.15fr); }
  .omics-ledger-action { grid-column: 1 / -1; justify-self: end; }
  .catalogue-filter-grid { grid-template-columns: minmax(300px, 1fr) repeat(2, minmax(150px, .45fr)) auto; }
  .catalogue-filter-grid > label:nth-of-type(3) { grid-column: 2; }
  .genome-view-panel[data-genome-view="circular"] { grid-template-columns: minmax(0, 1fr) 270px; }
  .genome-support-grid { grid-template-columns: minmax(0, 1fr) 270px; }
  .accession-chip-ledger { grid-template-columns: repeat(2, minmax(0, 1fr)); }
}

@media (max-width: 980px) {
  .species-record-identity,
  .gene-record-identity,
  .research-record-heading,
  .record-identity.accession-record-identity { grid-template-columns: 1fr; align-items: start; }
  .species-record-actions { justify-content: flex-start; }
  .record-layout,
  .gene-record-layout,
  .genome-support-grid { grid-template-columns: 1fr; }
  .record-aside-unified { position: static; grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .record-aside-unified > .alert,
  .record-aside-unified > .status-detail { grid-column: 1 / -1; }
  .species-overview-grid { grid-template-columns: 1fr; }
  .species-portrait-panel { min-height: 320px; }
  .evidence-ribbon-grid { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .omics-ledger-row { grid-template-columns: minmax(240px, 1fr) minmax(180px, .65fr); }
  .omics-ledger-facts { grid-column: 1 / -1; }
  .omics-ledger-action { grid-column: auto; align-self: center; }
  .genome-view-panel[data-genome-view="circular"] { grid-template-columns: 1fr; }
  .complete-inspector { border-left: 0; border-top: 1px solid var(--line-strong); }
  .expression-viewer-stage { grid-template-columns: 1fr; }
  .heatmap-inspector-complete { border-left: 0; border-top: 1px solid var(--line-strong); }
  .catalogue-filter-grid { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .catalogue-filter-grid .filter-search { grid-column: 1 / -1; }
  .taxonomy-ledger { grid-template-columns: 1fr; }
  .organelle-layer-entry { grid-template-columns: 120px minmax(0, 1fr) 20px; padding: 18px; }
  .organelle-layer-art { width: 110px; height: 110px; }
  .source-ledger { grid-template-columns: 1fr; }
}

@media (max-width: 760px) {
  .complete-record-content { padding-top: 18px; }
  .species-record-identity,
  .gene-record-identity,
  .research-record-heading,
  .record-identity.accession-record-identity { gap: 16px; padding: 17px 0 20px; }
  .species-record-identity h1,
  .gene-record-identity h1,
  .research-record-heading h1 { font-size: clamp(34px, 10vw, 45px); }
  .gene-full-name { font-size: 18px; }
  .species-record-actions,
  .page-actions { width: 100%; display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .species-record-actions .button:first-child:nth-last-child(3) { grid-column: 1 / -1; }
  .page-actions .button { width: 100%; }
  .species-context-navigation .context-navigation-inner,
  .gene-context-navigation .context-navigation-inner { gap: 20px; }
  .evidence-ribbon { padding: 14px; }
  .evidence-ribbon-grid { grid-template-columns: 1fr; border-left: 0; border-top: 1px solid var(--line); }
  .evidence-layer-cell { border-left: 1px solid var(--line); }
  .record-aside-unified { grid-template-columns: 1fr; }
  .section-block + .section-block { margin-top: 34px; }
  .section-heading { align-items: flex-start; flex-direction: column; gap: 8px; }
  .section-heading h2 { font-size: 22px; }
  .editorial-note-list article,
  .record-prose-facts div { grid-template-columns: 1fr; gap: 5px; }
  .species-portrait-panel img { min-height: 280px; }
  .sequence-record-row { grid-template-columns: minmax(0, 1fr) auto; }
  .sequence-record-row code { grid-column: 1; }
  .pathway-record-summary { grid-template-columns: 1fr; }
  .cross-layer-grid,
  .record-stat-grid,
  .assembly-summary-grid { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .cross-layer-stat:nth-child(2),
  .record-stat-grid > div:nth-child(2),
  .assembly-metric:nth-child(2) { border-right: 0; }
  .cross-layer-stat:nth-child(-n+2),
  .record-stat-grid > div:nth-child(-n+2),
  .assembly-metric:nth-child(-n+2) { border-bottom: 1px solid var(--line); }
  .research-path-list { grid-template-columns: 1fr; }
  .omics-ledger-row { grid-template-columns: 1fr; gap: 12px; }
  .omics-ledger-facts { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .omics-ledger-facts div { border-left: 0; border-top: 1px solid var(--line); padding: 8px 0; }
  .omics-ledger-action { justify-self: stretch; }
  .omics-ledger-action .button { width: 100%; }
  .research-toolbar { overflow-x: auto; }
  .research-toolbar-inner { width: max-content; min-width: 100%; padding-inline: 8px; }
  .research-toolbar-title { min-width: 150px; }
  .research-toolbar-search { min-width: 240px; flex-basis: 240px; }
  .research-toolbar .button span { display: none; }
  .genome-canvas.complete-genome-canvas { min-height: 430px; padding: 4px; }
  .complete-genome-canvas .organelle-map-svg { width: 790px; max-width: none; }
  .complete-inspector { min-height: 260px; }
  .linear-genome-complete { width: 1120px; padding-inline: 34px; }
  .feature-table-wrap { max-height: 520px; }
  .expression-summary-strip { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .expression-summary-strip > div:nth-child(2) { border-right: 0; }
  .expression-summary-strip > div:nth-child(-n+2) { border-bottom: 1px solid var(--line); }
  .expression-matrix-complete { grid-template-columns: 140px repeat(var(--heatmap-columns), 56px); min-width: calc(140px + (var(--heatmap-columns) * 56px)); }
  .heatmap-corner,
  .heatmap-row-label { width: 140px; }
  .expression-legend-complete { align-items: flex-start; flex-direction: column; gap: 8px; }
  .sequential-legend i { width: 190px; }
  .structured-metadata-grid { grid-template-columns: 1fr; }
  .catalogue-toolbar { align-items: stretch; }
  .catalogue-search { flex-basis: 100%; }
  .compact-select { width: 100%; }
  .catalogue-filter-grid,
  .accession-filters { grid-template-columns: 1fr; }
  .catalogue-filter-grid .filter-search,
  .catalogue-filter-grid > label:nth-of-type(3) { grid-column: auto; }
  .catalogue-summary-row { align-items: flex-start; flex-direction: column; gap: 3px; }
  .table-wrap { max-width: 100%; overflow-x: auto; }
  .scientific-table { min-width: 760px; }
  .scientific-table .sticky-column { max-width: 180px; }
  .gene-sequence-ledger article { grid-template-columns: 1fr; gap: 8px; }
  .gene-sequence-ledger pre { white-space: pre; overflow-x: auto; }
  .accession-chip-ledger { grid-template-columns: 1fr; }
  .related-gene-list a { grid-template-columns: 76px minmax(0, 1fr) 18px; }
  .related-gene-list small { grid-column: 2; }
  .evidence-definition-row,
  .organelle-layer-split { grid-template-columns: 1fr; }
  .organelle-layer-entry { min-height: 190px; grid-template-columns: 100px minmax(0, 1fr) 18px; }
  .organelle-layer-art { width: 90px; height: 90px; }
  .taxonomy-ledger > a { padding: 13px; }
  .reference-ledger article { grid-template-columns: 45px minmax(0, 1fr); }
  .reference-ledger .button { grid-column: 2; justify-self: start; }
  .source-ledger a { grid-template-columns: minmax(80px, .4fr) minmax(0, 1fr) 16px; }
  .gene-evidence-strip,
  .record-meta-strip { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .gene-evidence-strip > div,
  .record-meta-strip > div { border-bottom: 1px solid var(--line); }
  .gene-evidence-strip > div:nth-child(2n),
  .record-meta-strip > div:nth-child(2n) { border-right: 0; }
  .gene-evidence-strip > div:last-child,
  .record-meta-strip > div:last-child { grid-column: 1 / -1; border-bottom: 0; }
}

@media (max-width: 420px) {
  .species-record-actions,
  .page-actions { grid-template-columns: 1fr; }
  .species-record-actions .button:first-child:nth-last-child(3) { grid-column: auto; }
  .cross-layer-grid,
  .record-stat-grid,
  .assembly-summary-grid { grid-template-columns: 1fr; }
  .cross-layer-stat,
  .record-stat-grid > div,
  .assembly-metric { border-right: 0; border-bottom: 1px solid var(--line); }
  .cross-layer-stat:last-child,
  .record-stat-grid > div:last-child,
  .assembly-metric:last-child { border-bottom: 0; }
  .omics-ledger-facts { grid-template-columns: 1fr; }
  .expression-summary-strip { grid-template-columns: 1fr; }
  .expression-summary-strip > div { border-right: 0; border-bottom: 1px solid var(--line); }
  .expression-summary-strip > div:last-child { border-bottom: 0; }
  .organelle-layer-entry { grid-template-columns: 70px minmax(0, 1fr) 16px; gap: 12px; padding: 14px; }
  .organelle-layer-art { width: 65px; height: 65px; }
  .organelle-layer-art svg { width: 31px; height: 31px; }
}

/* Global multi-omics entry ------------------------------------------------- */
.omics-entry-page { background: linear-gradient(180deg, color-mix(in srgb, var(--lavender-75) 42%, var(--page)) 0, var(--page) 340px); }
.omics-entry-heading { padding-bottom: 28px; }
.omics-entry-index { border-block: 1px solid var(--line-gold); background: color-mix(in srgb, var(--surface) 92%, var(--lavender-75)); }
.omics-entry-index-inner { display: grid; grid-template-columns: repeat(5, minmax(0, 1fr)); }
.omics-entry-index a { min-width: 0; min-height: 82px; display: grid; grid-template-columns: 38px minmax(0, 1fr); align-items: center; gap: 11px; padding: 13px 16px; color: var(--ink); border-right: 1px solid var(--line); }
.omics-entry-index a:first-child { border-left: 1px solid var(--line); }
.omics-entry-index a:hover { background: var(--surface-raised); }
.omics-entry-index svg { width: 34px; height: 34px; color: var(--gold-700); }
.omics-entry-index strong,
.omics-entry-index small { display: block; }
.omics-entry-index strong { font-family: var(--font-editorial); font-size: 15px; }
.omics-entry-index small { margin-top: 2px; color: var(--muted); font-size: 10px; line-height: 1.3; }

.omics-entry-layer-ledger { border-top: 1px solid var(--line-strong); }
.omics-entry-layer { display: grid; grid-template-columns: minmax(0, 1fr) 300px; align-items: stretch; border-bottom: 1px solid var(--line-strong); }
.omics-entry-layer-main { min-width: 0; display: grid; grid-template-columns: 64px minmax(0, 1fr) 22px; align-items: center; gap: 16px; padding: 20px 22px 20px 8px; color: var(--ink); }
.omics-entry-layer-main:hover { background: linear-gradient(90deg, transparent, var(--surface-soft)); }
.omics-entry-layer-icon { width: 60px; height: 60px; display: grid; place-items: center; color: var(--lavender-700); }
.omics-entry-layer-icon svg { width: 54px; height: 54px; }
.omics-entry-layer.is-chloroplast .omics-entry-layer-icon,
.omics-entry-layer.is-mitochondrial .omics-entry-layer-icon { color: var(--gold-700); }
.omics-entry-layer-copy strong { display: block; font-family: var(--font-editorial); font-size: 22px; }
.omics-entry-layer-copy small { display: block; max-width: 760px; margin-top: 5px; color: var(--muted); font-size: 13px; line-height: 1.52; }
.omics-entry-arrow { width: 20px; height: 20px; color: var(--gold-700); }
.omics-entry-layer-metrics { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); align-content: center; gap: 8px 14px; padding: 16px 0 16px 24px; border-left: 1px solid var(--line); }
.omics-entry-layer-metrics div { display: grid; gap: 2px; }
.omics-entry-layer-metrics span { color: var(--muted); font-size: 10px; text-transform: uppercase; letter-spacing: .06em; }
.omics-entry-layer-metrics strong { color: var(--ink); font-family: var(--font-editorial); font-size: 22px; font-variant-numeric: tabular-nums; }
.omics-entry-layer-metrics > a { grid-column: 1 / -1; color: var(--gold-700); font-size: 12px; font-weight: 600; }

.workflow-ledger { border-top: 1px solid var(--line-strong); }
.workflow-ledger > a { display: grid; grid-template-columns: 52px minmax(0, 1fr) 22px; align-items: center; gap: 18px; padding: 18px 8px; color: var(--ink); border-bottom: 1px solid var(--line-strong); }
.workflow-ledger > a:hover { background: linear-gradient(90deg, transparent, var(--surface-soft), transparent); }
.workflow-index { color: var(--gold-700); font-family: var(--font-mono); font-size: 13px; }
.workflow-ledger strong,
.workflow-ledger small { display: block; }
.workflow-ledger strong { font-family: var(--font-editorial); font-size: 18px; }
.workflow-ledger small { max-width: 880px; margin-top: 4px; color: var(--muted); line-height: 1.5; }
.workflow-ledger > a > svg { width: 19px; height: 19px; color: var(--gold-700); }
.workflow-ledger code { color: var(--ink); font-size: 12px; }

.coverage-track { width: min(240px, 100%); height: 6px; overflow: hidden; background: var(--neutral-100, var(--line)); border-radius: 2px; }
.coverage-track span { display: block; height: 100%; background: linear-gradient(90deg, var(--lavender-700), var(--gold-700)); }

.evidence-definition-ledger { display: grid; grid-template-columns: repeat(4, minmax(0, 1fr)); border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.evidence-definition-ledger article { min-height: 150px; padding: 20px; border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.evidence-definition-ledger p { margin: 15px 0 0; color: var(--muted); font-size: 13px; line-height: 1.55; }

.related-data-split { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); border-top: 1px solid var(--line); border-left: 1px solid var(--line); }
.related-data-split article { min-height: 210px; display: grid; grid-template-columns: 56px minmax(0, 1fr); gap: 18px; padding: 24px; border-right: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.related-data-split article > svg { width: 52px; height: 52px; color: var(--gold-700); }
.related-data-split h3 { margin: 0; font-family: var(--font-editorial); font-size: 21px; }
.related-data-split p { margin: 7px 0 18px; color: var(--muted); line-height: 1.56; }
.related-data-split .button-row { display: flex; align-items: center; flex-wrap: wrap; gap: 12px; }

html[data-theme="dark"] .omics-entry-index { background: color-mix(in srgb, var(--surface) 88%, var(--lavender-75)); }
html[data-theme="dark"] .omics-entry-page { background: linear-gradient(180deg, rgba(95, 64, 124, .12) 0, var(--page) 350px); }

@media (max-width: 1100px) {
  .omics-entry-index-inner { grid-template-columns: repeat(3, minmax(0, 1fr)); }
  .omics-entry-index a:nth-child(4), .omics-entry-index a:nth-child(5) { border-top: 1px solid var(--line); }
  .omics-entry-layer { grid-template-columns: minmax(0, 1fr) 250px; }
  .evidence-definition-ledger { grid-template-columns: repeat(2, minmax(0, 1fr)); }
}

@media (max-width: 760px) {
  .omics-entry-index-inner { grid-template-columns: 1fr; }
  .omics-entry-index a,
  .omics-entry-index a:first-child { min-height: 68px; border-left: 0; border-right: 0; border-top: 1px solid var(--line); }
  .omics-entry-index a:first-child { border-top: 0; }
  .omics-entry-layer { grid-template-columns: 1fr; }
  .omics-entry-layer-main { grid-template-columns: 48px minmax(0, 1fr) 18px; gap: 12px; padding: 17px 4px; }
  .omics-entry-layer-icon { width: 46px; height: 46px; }
  .omics-entry-layer-icon svg { width: 42px; height: 42px; }
  .omics-entry-layer-copy strong { font-size: 18px; }
  .omics-entry-layer-copy small { font-size: 11px; }
  .omics-entry-layer-metrics { grid-template-columns: repeat(3, minmax(0, 1fr)); padding: 13px 0; border-left: 0; border-top: 1px solid var(--line); }
  .omics-entry-layer-metrics > a { grid-column: auto; align-self: end; text-align: right; }
  .workflow-ledger > a { grid-template-columns: 35px minmax(0, 1fr) 18px; gap: 10px; padding-inline: 0; }
  .workflow-ledger strong { font-size: 16px; }
  .evidence-definition-ledger { grid-template-columns: 1fr; border-left: 0; }
  .evidence-definition-ledger article { min-height: auto; padding-inline: 0; border-right: 0; }
  .related-data-split { grid-template-columns: 1fr; border-left: 0; }
  .related-data-split article { min-height: auto; grid-template-columns: 44px minmax(0, 1fr); padding: 20px 0; border-right: 0; }
  .related-data-split article > svg { width: 40px; height: 40px; }
}